Yooka-Replaylee: Quills Locations & Interactive Tracking Guide

Found on top of the pillar. You can reach them using the Flight Bomb. After you grab it, MASH the fly button to ascend.

Found on the lower outcrop of the large stone. You can reach them by dropping from above.

Found on top of the green mushroom. Hop on it to collect the Quills.

Found behind a tree on a ledge.

Found next to the mountain wall, on a lower ledge.

Found between the icicles.

Found behind the large icicles.

Found at the bottom of the small pond.

Found on a pile of gold inside Rampo's Treasure Area. You can access it via the entrance under the temple.

Found inside the cave accessed via the southeastern tunnel entrance.

Found behind the large pipe.

Found on the upper balcony. Jump on the green mushroom below to reach them.

Found on top of the rock. You can reach them using the Flight Bomb on top of the temple. Grab the Bomb and MASH the fly button to get to the rocks. Land on them to get the Quills.

Found on a lily pad.

Found on a ledge halfway up the mountain wall.

Found on a patch of land under the large tree.

Found on the lower ledge around the tower. You can reach them by dropping from above.

Found underwater, between two wooden pillars.

Found behind some crates inside the cylinder. Use the flight bombs to ascend and get the quills.

Found underwater, between some rocks.

Found on top of the cylinder. Use the flight bombs to ascend and get the quills.

Found on the roof of the lighthouse cylinder. Grab the flight bomb on the ground floor inside the lighthouse, and use it to fly to the top.

Found by the water, on the left side of the cave entrance.

Found in the second room of the cave you can access via the pipe entrance near Clara. Use the Sonar 'Splosion to reveal them.

Found on the shore, by the lake.

Found behind the wooden crates at the end of the dock.

Found on top of the stone statue.

Found on top of a pipe on the upper platform at the end of the course starting behind Jack-O-Lantern. Step on the pressure plates and use the grappling points to ascend the trees.

Found between two pillars at the bottom of the lake.

Found between two pillars at the bottom of the lake.

Found on top of a pipe. You can get to the roof by stepping on the pressure plate at the edge of the plateau, thus revealing a grappling point. After you reach the roof, Spring Jump on top of the pipe.

Found in the caged area above Vendi, in the first room of the Icymetric Palace. Use the Sonar Shot to reveal the invisible moving platforms and hop on them to get to the Quills.

Found along the wall, in the second room of the Icymetric Palace. Use the Reptile Roll to quickly collect them.

Found along the bridge, in the third room of the Icymetric Palace.

Found on a balcony in the room where you find Burnie. You can collect them after you defeat Brrreeze Blok by gliding in the air stream.

Found in Rextro's room. You can collect them on your way back to the entrance after defeating Brrreeze Blok.

Found in the second room east of the second Icymetric Palace room. You can access it on your way back to the entrance after defeating Brrreeze Blok.

Found behind some crates in the corridor.

Found in the gauntlet room north of Vendi. You can access it after defeating the boss. Collect the Quills while working on completing the gauntlet, on your way to the Toxic Trouble Pagie.

Found on the die behind the arcade machine.

The quills are found in the corridor past the first room east of Keepy Uppy Pagie. The first room, however, features a small puzzle you must solve by going through the entrances in the correct order: South, South, East, North.

Found behind some breakable crates on the ground floor.

Found by the stairs leading to the art gallery.

Found in the flooded room of the Icymetric Palace. You can access it via the entrance at the base of the tower. Once you get inside, swim up and look for them under the wooden platform.

Found on the ground floor, behind the large pile of chips.

Found inside the third room of the Icymetric Palace when you access it via the underwater entrance at the base of the tower. After reaching the room, use the nearby bomb to destroy the rocks covering the alcoves.

Found behind the slots.

Found inside the fourth room of the Icymetric Palace when you access it via the underwater entrance at the base of the tower. After reaching the room, you'll see them around the tub.

Found on top of the gate. You can reach them by dropping from above.

Found in the breakable displays.

Found on the ground under the table.

Found in the back of the broken slot machine.

Found next to a palm tree by the water.

Found on the small island.

Found on the ground behind the large golden statue.

The Quills are located under the pier, and you can obtain them after you bring the Mollycool to Dr. Puzz so she can transform you into a ship. Use the canons to destroy the boulder.

Found under the ramp next to Mark.

Found in front of the large structure.

Found behind the roulette.

Found by the stairs.

Found behind the planters by the wall.

Found at the end of the maze. You can collect them after you bring the Mollycool to Dr. Puzz so she can turn you into a ship.

Found inside the large boulder you can destroy after you bring the Mollycool to Dr. Puzz, so she can turn into a ship.

Found on the dice near the roulettes.

Found on the upper floor of the casino, on the cube behind Mark.

Found at the end of the island.

Found on the roof of the room. You can reach them using the grappling points at the edge of the wall.

Found in the corner of the roof. Use the Spring Jump to ascend.

Found on top of Rextro's arcade machine.

Found in the back of the stone blocks.

Found at the edge of the roof.

Found in the corner of the roof.

Found at the bottom of the pond.

Found on top of the tower.

Found under the breakable crates in the fountain.

Found on top of the hill, behind Lady Leapalot.

Found behind the broken slot machine.

Found behind the breakable crates.

Found behind the planter.

Found behind the planter.

Found by the wall near the broken slot machine.

Found at the bottom of the pond inside the Vault.

Found under the palm trees.

Found on the small island behind the breakable crates.

Found at the base of the roof. You can reach it using the grappling points on the rocks on the western side.

Found at the edge of the plateau.

Found on the golf course you can reach via the elevator.

Found on the golf course, behind an outcrop.

Found in the underwater tunnel, in front of a locked gate.

Found behind a tree.

Found on the ground under a large mushroom.

Found under the dice wall.

Found on the stone wall, behind a mushroom.

Found on top of the rocks by the tree.

Found on the jib boom of the ship.

Found near the pile of dice and some wooden crates.

Found in the back of the ship.

Found halfway up the must. You can reach them by gliding into the airstream below.

Found on the wooden platform in the back of the arena.

Found on top of the pipe you can reach by gliding.

Found between some mushrooms in the back of the plateau.

Found on the upper floor of the Captain's Cabin, accessed via the door on the ship.

Found inside a breakable crate at the edge of the plateau.

Found at the edge of the lower plateau.

Found on the wooden platform under a large tree.

Found on a mushroom near a large tree.

Found in the back of the structure.

Found under the bridge.

Found next to the palm tree on the shore.

Found on the roof of the building.

Found behind the stones by the water.

Found on a pipe.

Found on top of the robot. Jump on the green mushrooms below to reach them.

Found on a rock in a small enclosure you can reach by pulling the crate blocking the entrance.

Found under the stone blocks by the mountain wall. You can destroy them using the bomb at the edge of the plateau.

Found on top of a large tank. You can reach it by gliding after collecting the Lash Chance Pagie.

Found on the robot's leg. Glide from the upper side to reach them.

Found in front of Kartos in the back of the island.

Found at the edge of the island.

Found in the clearing, between the large stone skulls.

Found between the large trees.

Found on the ground, next to Mark.

Found on the ledge behind Mark.

Found between 2 mushrooms.

Found on the stone blocks while climbing the tower.

Found next to Normy in the enclosure at the edge of the swamp. Jump on the floating logs to get to the enclosure.

Found at the edge of the plateau.

Found behind some breakable crates in the upper room you can reach after you give Dr. Puzz the Mollycool so she can transform you into a helicopter.

Found at the end of the plateau.

Found behind the breakable wooden crates.

Floating above the swamp. You can collect them while riding the floating logs.

Found at the edge of the roof. After you give Dr. Puzz the Mollycool, turn into a helicopter to reach the upper roof of the house of cards.

Found on the lower ledge by the river.

Found behind the large arcade machine.

Found in the back of the large arcade game.

Found at the edge of the roof. After you give Dr. Puzz the Mollycool, turn into a helicopter to reach the upper roof of the house of cards.

Found on top of a red die. After you give Dr. Puzz the Mollycool, turn into a helicopter to reach the upper roof of the house of cards.

Found on top of a tree above the water. Use the grappling point to climb.

Found on the platform you can reach after you complete Schell's mission.

Found at the bottom of the lake, under the large tree.

Found on the lower roof of the casino. After you give Dr. Puzz the Mollycool, turn into a helicopter to reach the upper side of the casino.

Found on the upper platform. After you give Dr. Puzz the Mollycool, turn into a helicopter to reach the upper side of the casino.

Found behind three mushrooms at the edge of the lake.

Found inside the cave, you can access via the entrance at the bottom of the large western trench.

Found at the edge of the middle roof. After you give Dr. Puzz the Mollycool, turn into a helicopter to reach the upper side of the casino.

Found on top of the broken pillar.

Found by the mountain wall.

Found at the bottom of the pond.

Found on top of the pipe.

Found next to a large rock at the edge of the plateau.

Found on the lower roof of the house of cards. After you give Dr. Puzz the Mollycool, turn into a helicopter to reach the roof.

Found behind a large rock by the mountain wall.

Found behind a barrel and a crate by the mountain wall.

Found at the bottom of the lake, under the Pagie. To collect the Quills, you have to bring the Mollycool to Dr. Puzz, so she can transform you into a shoal. Once transformed, attack the thorns, collect the Pagie and the Quills.

Found on top of the large stone. You can reach them by flying after you grab the bomb in the outhouse.

Found on a rock by a stone pillar.

Found behind a tree at the end of the hill path.

Found behind a pillar on the upper side of the mountain. You can reach this area by stepping on the pressure plate in the southern clearing.

Found at the end of the path halfway up the mountain.

Found on a ledge next to Vendi.

Found underwater alongside 2 chests full of Q.U.I.D.S.

Found on a ledge.

Found on top of the golf flag. After you give Dr. Puzz the Mollycool, turn into a helicopter to reach the Quills.

Found behind a stone on the northern side of the mountain.

Found in front of the upper tunnel entrance.

Found on a ledge above Blastooie. You can reach it via the invisible platform near the cannon.

Found at the bottom of the frozen pond inside the cave. To melt the ice, set the four braziers in the cave on fire.

Found on a ledge under the wooden bridge.

Found on the upper ledge. After you give Dr. Puzz the Mollycool, turn into a helicopter to reach the Quills.

Found on the path outside the monument.

Found by the wall inside the cave.

Found behind some breakable crates.

Found on the balcony in front of the tunnel entrance.

Found on the left side of the ramp. To collect them, consider using the heavy bomb by the western entrance.

Found at the edge of the plateau.

Found inside a stone block you can destroy using the Throwing Bomb nearby.

Found under the wooden bridge.

Found behind the wooden gate, you can lower by slamming the pressure plate.

Found behind some breakable crates.

Found under the stone bridge.

Found on top of an ice block inside the tower.

Found on the wall.

Found outside the tower.

Found by the wall on top of the temple.

Found on top of a ledge. You can reach it by gliding from the southern tower platforms.

Found on the ground behind some breakable crates.

Found on top of a rock, by the large stone wall.

Found on top of the temple, behind Rampo. You can obtain them after defeating the boss.

Found on top of the dice pile. You can reach them by Spring Jumping on the edges of the dice.

Found under the bridge.

Found on the ground in the back of the cave.

Found at the bottom of the lake under an icy outcrop.

Found at the bottom of the lake.

Found at the bottom of the lake.

Found at the edge of the plateau.

Found next to the pipe entrance.

Found by the tree.

Found between the slippery rocks.

Found behind the rocks.

Found on the ground by the temple's wall.

Found behind a tree by the stone wall.

Found on top of a stone pillar.

Found on the ground, behind the tree.

Found on the lower ledge of the mushroom plateau.

Found on the mushroom in the swamp.

Found behind a rock by the swamp.

Found on top of the stone pillar.

Found by the pipe.,

Found on top of the stone pillar.

Found at the edge of the central plateau.

Found at the edge of the island, behind Vendi.